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P2, I think his mistake was worse than P1's because lowish middle set is even more vulnerable to a suck out in a 4 way pot.
No, his mistake (and it is the worst in the hand) is because of lost value, not because of vulnerability. He loses the chance to get stacks in vs TP by skipping the flop bet.
Next worse was P4's first turn call. He was far from priced in. He had an 8 out draw, good for 16% equity by the "rule of 2 and 4" estimate and 18% by the actual math. The first call is particularly bad, as he has no way of knowing P1 has anything and P2 could be stabbing or betting a weak made hand against which P4 would have no IOs OTR if his draw came in. That call of $75 represented 31.65% of the pot, WAY more than his outs justify. The second call at least could make an IOs argument, since by then it is clear that one or both of P1 and P2 have a hand they will pay off a river bet with. He's only paying a couple percent more than his direct odds, so the call is fine with good expectation if he hits.
Third worst was P2's turn call, as I discussed above, as it basically forced the river call.
P1 played it fine, imo. I would mildly prefer a bet OTF, just because it is so multi-way and we want to build the pot, but I understand her thought that she crushed the board so hard that she wouldn't get value.